Carmen Marie Greenwood

OLYMPUS DIGITAL CAMERATitle: Assistant Professor

Department: Fisheries, Wildlife and Environmental Studies


Office Location: 127 CANR

Office Phone: 518-255-5584

Areas of Expertise: Entomology, Conservation Biology, Disturbance Ecology, Terrestrial Invertebrate Ecology

Courses Taught:

  • FWLD 101 Introduction to Natural Resource Conservation
  • BIOL 186 Entomology
  • AGRN 121 Soil and Water Conservation
  • BIOL 308 Terrestrial Invertebrate Ecology
  • ENVR 325 Research methods I
  • ENVR 326 Research methods II
  • BIOL 403 Conservation Biology

Curriculum Vitae >

About Me:

I am a broadly trained invertebrate ecologist and conservation biologist. Students in my lab have addressed questions related to the impacts of disturbance on invertebrate communities, food web interactions (such as forage availability for vertebrate wildlife), and conservation. Disturbance may occur naturally in a system or result from human activities such as tillage, prescribed burning, grazing, soil amendment, development, habitat fragmentation, compaction or invasive plant species. We are currently heavily involved in research focused on conservation of the federally endangered American Burying Beetle. If you are interested in invertebrate field ecology or behavioral ecology and would like to join our very “student-focused” research team please email me regarding your specific interests.

Projects currently underway in our lab:

  1. Expanded methodologies for long term monitoring of relocation survival and population density of federally endangered American Burying Beetle (Nicrophorus americanus) in Oklahoma, and surveys for potential re-introduction in central New York. ( duration Sep 2012- Dec 2015). Student researchers involved:
    a. Greg Strait, B.T. Wildlife Management SUNY Cobleskill
    b. Ben Derr, B.T. Wildlife Management SUNY Cobleskill
    c. Thomas Ferrari, M.S. candidate in Entomology and Plant Pathology, OSU
    d. Kyle Risser, Ph.D. candidate in Entomology and Plant Pathology, OSU
  2. The Effect of Preferred Arthropod Availability on Bobwhite Quail Nest Site Selection and Chick Survival (duration: July 2011- Dec 2014): This is a large, interdisciplinary project involving multiple collaborators and multiple states. Graduate students working on this project are evaluating the effects of a variety of factors that impact arthropod community composition within Beaver and Packsaddle Wildlife management areas of western Oklahoma. Studies focus on arthropod taxa known to be key forage species for juvenile quail. Oklahoma Dept. of Wildlife Conservation. Student researchers involved (from left to right above):
    a. Kenneth Masloski, M.S. Entomology and Plant Pathology: Community composition and resource partitioning of Short-horned grasshoppers (Orthoptera: Acrididae) in western Oklahoma grassland ecosystems.
    b. Allison Giguere; M.S. candidate in Entomology and Plant Pathology: Community composition and resource partitioning of ants (Hymenoptera: Formicidae) in western Oklahoma grassland ecosystems.
    c. Shane Foye; M.S. candidate in Entomology and Plant Pathology: Characterization of ground-dwelling arthropod assemblages and Entomopathogenic nematodes in Northern bobwhite quail (Colinus virginianus) habitat.

Projects recently completed with students:

  1. Profile of ecoregion and management practices impacts on prevalence of native entomopathogenic nematodes, and community composition of soil-dwelling microarthropods (primarily soil mites) in Oklahoma. Student researchers involved:
    a. Kyle Risser; M.S. Entomology and Plant Pathology, OSU
  2. Alexandra Robideau; undergraduate Niblack scholar (2011) in Entomology and Plant Pathology: Prevalence of native Entomopathogenic nematodes (EPN) in organic versus conventional wheat and beef production systems in Oklahoma
  3. Mackenzie Jochim; undergraduate Niblack scholar (2012) in Entomology and Plant Pathology: Impacts of Grazing (Cattle vs. Bison) and Controlled Burning on Entomopathogenic Nematode (EPN) Prevalence in the Tallgrass Prairie Preserve
  4. Colby Gregg: undergraduate Niblack scholar (2012) in Entomology and Plant Pathology: Native pollinator conservation and biological monitoring of the Cow Creek remediation project at IERES (Integrated Environmental Research and Education Site)


Refereed publications (*publications with students):

*Foye, S., C. Greenwood, M. Payton and K. Masloski. 2014. Characterization of ground-dwelling arthropod forage communities that support Colinus virginianus (Galliformes: Odontophoridae) in two western Oklahoma Wildlife Management Areas. Rangeland Ecology and Management. In preparation.

*Foye, S., C. Greenwood, and M. Payton. 2014. Virulence of indigenous entomopathogenic nematodes (EPN) towards Diorhabda carinulata, a potential biological control agent for the reduction of Tamarix ramosissima in northern bobwhite habitat. Southwestern Entomology. In preparation

*Giguere, A., C. Greenwood, and M. Payton. 2014. Community composition and resource partitioning of ants (Hymenoptera: Formicidae) in western Oklahoma grassland ecosystems: a critical forage taxon of the northern bobwhite (Colinus virginianus). Rangeland Ecology and Management. In preparation.

Greenwood, C., and P. Gagnon. 2014. Preliminary survey of carrion beetle (Coleoptera: Silphidae) community composition in the Land Between the Lakes National Recreation Area of Western Kentucky: A site targeted for re-introduction of American Burying Beetle (Nicrophorus americanus). Conservation Biology. In preparation.

*Masloski, K., C. Greenwood, M. Payton. 2014. Grasshopper (Orthopter: Acrididae) Communities in Northern Bobwhite (Colinus virginianus) habitat: spatial and temporal prey availability in a Western Oklahoma Grassland. Journal of Conservation Biology. In review

*Masloski, K., C. Greenwood, M. Payton, M. Reiskind. 2014.Grasshopper (Orthoptera: Acrididae) relative abundance and density: a comparison between standard and novel methods of sampling. Journal of Orthoptera Research. Accepted.

*Masloski, K., M. Payton, M. Reiskind, C. Greenwood. 2014. Evidence for diet-driven habitat partitioning of Melanoplinae and Gomphocerinae grasshoppers (Orthoptera: Acrididae) along a vegetation gradient in a Western Oklahoma grassland. Environmental Entomology. Accepted.
Antonenko, P. and C. Greenwood. 2014. Fostering collaborative problem solving and 21st century skills using the DEEPER scaffolding framework. Journal of College Science Teaching. Accepted.

*Risser, K. and C. Greenwood. 2014. Entomopathogenic nematode (EPN) prevalence and diversity across a state wide precipitation gradient in Oklahoma. Southwestern Entomologist. In review

*Jochim, M., C. Greenwood and K. Risser. 2014. Impacts of grazing (Cattle vs. Bison) and controlled burning on Entomopathogenic Nematode (EPN) prevalence in the Tallgrass Prairie Preserve. Southwestern Entomologist. In preparation.

*Robideau, A., K. Risser and C. Greenwood. 2014. Prevalence of native entomopathogenic nematodes (EPN) in organic versus conventional wheat and beef production systems in Oklahoma. Southwestern Entomologist. In review.

*Masloski, K. and C. Greenwood. 2013. First record of Myrmecophilus nebrascensis (Orthoptera: Myrmecophilidae) in Beaver Co., Oklahoma. Journal of Orthoptera Research. 22(1): 69-71.

*Booher, E., C. Greenwood and J. Hattey. 2012. Effects of Soil Amendments on Soil Microarthropods in Continuous Maize in Western Oklahoma. Southwestern Entomologist. Vol. 37, No. 1, pp. 12-20.

*Jones, M. E., Antonenko, P. D., and C. Greenwood. 2012. The impact of collaborative and individualized Student Response System strategies on learner motivation, metacognition, and knowledge transfer. Journal of Computer Assisted Learning. 28(3): 477-487.

*Dubie, T. and C. Greenwood, C. Godsey and M. Payton. 2011. Effects of tillage on soil microarthropods in winter wheat. Southwestern Entomologist. 36(1): 11-21.

*Ibrahim, M., Antonenko, P., Greenwood, C., and Wheeler, D. 2011. Effects of segmenting, signaling, and weeding on learning from educational video. Learning, Media, and Technology. 37(3): 220-235.

Greenwood, C., M. Barbercheck and C. Brownie. 2011. Short term response of soil microinvertebrates to application of entomopathogenic nematode-infected insects in two tillage systems. Pedobiologia. (54)(3): 177-186.
Greenwood, C.M. and E.G. Maurakis. 1998. Breeding Behaviors in Notropis alborus (Actinopterygii: Cyprinidae). VA Journal of Science 49 (3): 163 -172.


Research and Grants:

Extramural Competitive Grants: Funded
Kentucky Academy of Science. 2014 $11,600. Occurrence of mammalian prey and scavengers on potential re-introduction sites for Nicrophorus americanus at Land Between the Lakes NRA. T. Derting, C. Greenwood, P. Gagnon, H. Passmore, J. Hardin.

Oklahoma Department of Wildlife Conservation – 2014-2016. $89,936. Assessment of the density and spatial and temporal variation of the American Burying Beetle (Nicrophorus americanus) in Oklahoma. Jan 2014- Dec 2016. B. Luttbeg, C. Greenwood, J. Hardin, K. Giles, S. McMurry.

Oklahoma Department of Wildlife Conservation – 2012-2015. $3,700,000 ($502,094 dedicated to Entomology portion of study with Dr. Greenwood as the PI of that section).Evaluation of Northern Bobwhite in Western Oklahoma. Aug 2011-July 2017. D. Leslie, C. Davis, D. Elmore, S. Fuhlendorf, F. Guthery, T. O’Connell, C. Greenwood.

XTO Energy Corp./U.S. Fish and Wildlife Service Sep 2012 – Dec 2015– . $1,130,000. Expanded methodologies for long term monitoring of American Burying Beetle (Nicrophorus Americana) survival and characterization of habitat suitability following relocation in Oklahoma. C. Greenwood, K. Giles, J. Hardin, B. Luttbeg and S. McMurray.

USDA Sustainable Agriculture Research and Education (SARE) graduate student award – 2000-2002. $10,000. Short term response of microinvertebrates to application of entomopathogenic nematode infected insect cadavers. C. Greenwood and M. Barbercheck.

Competitive Student Research Awards: Funded
McNair undergraduate research scholarship. January, 2014. Characterization of cuticular hydrocarbons of Nicrophorus orbicollis and N. pustulatus in the context of solitary versus communal mating. Nick Blumenberg and Carmen Greenwood

National Science Foundation Graduate Student Research Fellowship. March 2013 Impact of Diorhabda carinulata (Coleoptera: Chrysomelidae), an introduced biological control agent of Tamarix spp. on arthropod forage taxa in arid grasslands. Sunny Evans, M.S. Dept. of Entomology and Plant Pathology, Oklahoma State University.

Niblack undergraduate research scholarship. April, 2012. Conservation of native pollinators using indigenous wildflowers in riparian and wetland developments. Colby Gregg and Carmen Greenwood.

Niblack undergraduate research scholarship. April, 2011. Entomopathogenic nematode diversity of the Tallgrass prairie. Mackenzie Jochim and Carmen Greenwood.

Niblack undergraduate research scholarship. April, 2010. Naturally-occurring biological control of insect pests in organic wheat/beef systems. Alexandra Robideaux and Carmen Greenwood.
Wentz undergraduate research scholarship. April, 2010. Using soil mites as indicators of soil amendment efficacy. Evan Booher and Carmen Greenwood.

Johnston Award Scholarship – Ohio State University summer course in mite taxonomy. June, 2008. $500 scholarship awarded to M.S. student Trisha Dubie.


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